We use cookies to enhance the usability of our website. If you continue, we'll assume that you are happy to receive all cookies. More information. Don't show this again.
On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
8.4
nTPM
HPA:
5.5
nTPM
GTEx:
8.4
nTPM
FANTOM5:
10.5
Scaled Tags Per Million
PROSTATE - Annotated protein expression
Pending normal tissue analysis
PROSTATE - Antibody stainingi
Antibody staining in the annotated cell types in the current human tissue is reported as not detected, low, medium, or high, based on conventional immunohistochemistry profiling in selected tissues. This score is based on the combination of the staining intensity and fraction of stained cells.
Each image is clickable and will lead to virtual microscopy that enables deeper exploration of all samples and also displays staining intensity scores, fraction scores and subcellular localization as well as patient and tissue information for each sample.
Antibody HPA049807
Glandular cells
Not detected
PROSTATE - HPA RNA-seqi
The RNA-seq details section shows detailed information about the individual samples used for the transcript profiling and results of the RNA-seq analysis.
Information about each individual sample is listed below, including gender, age, a tissue section image and estimated fractions of cell types. nTPM (normalized transcripts per million) values give a quantification of the gene abundance which is comparable between different genes and samples.
Average nTPM
5.5
Male, age 62
Prostate sample 59 nTPM: 4.5
Cell types%
Glandular cells:
30
Smooth muscle cells:
55
Skeletal muscle cells:
5
Other cell types:
10
Male, age 65
Prostate sample 205 nTPM: 6.3
Cell types%
Glandular cells:
20
Smooth muscle cells:
60
Other cell types:
20
Male, age 65
Prostate sample 206 nTPM: 7.5
Cell types%
Glandular cells:
30
Smooth muscle cells:
65
Other cell types:
5
Male, age 68
Prostate sample 207 nTPM: 5.7
Cell types%
Glandular cells:
30
Smooth muscle cells:
65
Other cell types:
5
Male, age 60
Prostate sample 361 nTPM: 5.1
Cell types%
Glandular cells:
40
Smooth muscle cells:
55
Other cell types:
5
Male, age 66
Prostate sample 364 nTPM: 5.3
Cell types%
Glandular cells:
50
Smooth muscle cells:
45
Other cell types:
5
Male, age 63
Prostate sample 365 nTPM: 4.7
Cell types%
Glandular cells:
60
Smooth muscle cells:
35
Other cell types:
5
Male, age 66
Prostate sample 388 nTPM: 6.7
Cell types%
Glandular cells:
65
Smooth muscle cells:
30
Other cell types:
5
Male, age 58
Prostate sample 412 nTPM: 3.9
Cell types%
Glandular cells:
45
Smooth muscle cells:
50
Other cell types:
5
PROSTATE - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Average nTPM
8.4
GTEx sample id
Sample description
nTPM
Prostate Average nTPM: 8.4
Samples: 245
Max nTPM: 15.9
Min nTPM: 3.8
Std nTPM: 2.0
Median nTPM: 8.3
GTEX-1A3MV-0826-SM-72D6J
30-39 years, male
15.9
GTEX-1PPH8-1426-SM-DTXA3
20-29 years, male
13.4
GTEX-XQ8I-2526-SM-EAZB3
50-59 years, male
13.2
GTEX-RWSA-2026-SM-47JX8
40-49 years, male
13.1
GTEX-11OF3-1726-SM-5GU5Q
60-69 years, male
12.7
GTEX-WOFM-0326-SM-4OOSL
20-29 years, male
12.6
GTEX-Y9LG-1826-SM-4VBQC
30-39 years, male
12.6
GTEX-144GM-0826-SM-5O98R
20-29 years, male
12.4
GTEX-ZZ64-0926-SM-5E44L
20-29 years, male
12.4
GTEX-1PIGE-1726-SM-E6CQH
20-29 years, male
12.4
GTEX-17MFQ-1326-SM-718BX
20-29 years, male
12.2
GTEX-1QCLY-1826-SM-DTX89
30-39 years, male
12.2
GTEX-1QP29-0526-SM-EAZ3T
50-59 years, male
12.2
GTEX-1LBAC-1926-SM-DHXK2
20-29 years, male
12.1
GTEX-18QFQ-2026-SM-72D7J
30-39 years, male
12.0
GTEX-1KANB-2026-SM-DIPEY
50-59 years, male
12.0
GTEX-X5EB-1826-SM-4E3K8
40-49 years, male
11.9
GTEX-ZUA1-2826-SM-59HLE
40-49 years, male
11.8
GTEX-145LT-0626-SM-5O99E
40-49 years, male
11.5
GTEX-1C64O-2026-SM-7IGPA
60-69 years, male
11.5
GTEX-U4B1-1426-SM-4DXTX
40-49 years, male
11.2
GTEX-1I1GP-2026-SM-CE6SZ
50-59 years, male
11.1
GTEX-1N2EE-1026-SM-EXUS8
60-69 years, male
11.1
GTEX-QV44-1526-SM-DHXIP
50-59 years, male
11.1
GTEX-S4Q7-1026-SM-4AD75
20-29 years, male
11.0
GTEX-Y8E4-2126-SM-5RQHO
40-49 years, male
11.0
GTEX-YF7O-1926-SM-4W1YQ
50-59 years, male
11.0
GTEX-16YQH-2326-SM-7EWDS
30-39 years, male
10.9
GTEX-T8EM-1826-SM-4DM7F
40-49 years, male
10.9
GTEX-1CAMQ-2226-SM-7MGWS
40-49 years, male
10.9
GTEX-1QP2A-1526-SM-E6CPN
50-59 years, male
10.9
GTEX-18A66-2326-SM-7LT8Q
60-69 years, male
10.8
GTEX-ZDYS-1226-SM-5IJF3
50-59 years, male
10.8
GTEX-1I1GV-1926-SM-CNPQH
60-69 years, male
10.8
GTEX-12C56-1326-SM-5FQSV
20-29 years, male
10.7
GTEX-14PJO-2826-SM-6AJA6
60-69 years, male
10.7
GTEX-PLZ6-1126-SM-3P5ZR
30-39 years, male
10.5
GTEX-V1D1-1926-SM-4JBGX
50-59 years, male
10.5
GTEX-QV31-1026-SM-CNNOS
30-39 years, male
10.5
GTEX-148VJ-1826-SM-5Q5DV
70-79 years, male
10.4
GTEX-1CAMR-1626-SM-79OLC
20-29 years, male
10.4
GTEX-17EUY-2026-SM-79OMV
20-29 years, male
10.3
GTEX-11DYG-2526-SM-5N9BB
60-69 years, male
10.2
GTEX-132NY-2826-SM-5P9G2
60-69 years, male
10.2
GTEX-1I1GU-1126-SM-A96S9
40-49 years, male
10.2
GTEX-1RAZQ-0826-SM-EVR4L
30-39 years, male
10.2
GTEX-WFG8-1826-SM-4LVM4
20-29 years, male
10.1
GTEX-ZY6K-1526-SM-5GZXE
50-59 years, male
10.1
GTEX-111FC-2026-SM-5GZYO
60-69 years, male
10.0
GTEX-11TUW-2126-SM-5GU7B
60-69 years, male
10.0
GTEX-YB5E-1826-SM-5IFI4
40-49 years, male
10.0
GTEX-YJ8A-1126-SM-5IFJU
20-29 years, male
10.0
GTEX-1LG7Z-0826-SM-CXKYN
30-39 years, male
10.0
GTEX-Q2AH-1426-SM-EXOHC
40-49 years, male
10.0
GTEX-111CU-1526-SM-5N9FS
50-59 years, male
9.9
GTEX-13G51-2326-SM-5LZXV
60-69 years, male
9.9
GTEX-1JMQK-2226-SM-ARU8U
60-69 years, male
9.9
GTEX-1MUQO-1526-SM-DPRYK
60-69 years, male
9.9
GTEX-145LV-1226-SM-5S2QH
50-59 years, male
9.8
GTEX-1CB4F-2326-SM-7MKFO
60-69 years, male
9.8
GTEX-WZTO-0926-SM-4PQZ3
40-49 years, male
9.8
GTEX-1H1E6-0526-SM-9WPQ3
30-39 years, male
9.8
GTEX-1IL2U-2826-SM-A96SR
60-69 years, male
9.8
GTEX-P4PQ-2026-SM-E9U4E
60-69 years, male
9.8
GTEX-11NSD-0826-SM-5986S
20-29 years, male
9.7
GTEX-S33H-1826-SM-4AD65
30-39 years, male
9.7
GTEX-U412-2026-SM-4DXSI
60-69 years, male
9.7
GTEX-1I6K6-2126-SM-B2LX6
30-39 years, male
9.7
GTEX-117YX-1526-SM-5H12T
50-59 years, male
9.6
GTEX-1B97I-0826-SM-731DN
40-49 years, male
9.6
GTEX-NFK9-2226-SM-3MJGP
40-49 years, male
9.6
GTEX-OIZH-2026-SM-3NB1M
50-59 years, male
9.6
GTEX-WYJK-0626-SM-4ONCZ
50-59 years, male
9.6
GTEX-1P4AB-1926-SM-E6CQL
50-59 years, male
9.6
GTEX-1399R-0526-SM-5IJEF
30-39 years, male
9.5
GTEX-S95S-1026-SM-4B64M
60-69 years, male
9.5
GTEX-WH7G-1726-SM-4LVKY
40-49 years, male
9.5
GTEX-12WSN-5007-SM-7MGWH
40-49 years, male
9.5
GTEX-1CB4J-1526-SM-7MXVB
60-69 years, male
9.5
GTEX-1HBPI-0326-SM-A9G3N
50-59 years, male
9.5
GTEX-13FXS-2826-SM-5LZYC
50-59 years, male
9.4
GTEX-SNMC-0926-SM-4DM5U
20-29 years, male
9.4
GTEX-ZPU1-2026-SM-57WFI
40-49 years, male
9.4
GTEX-139T6-1526-SM-5P9G6
50-59 years, male
9.3
GTEX-13FHO-2826-SM-5K7ZA
60-69 years, male
9.3
GTEX-U3ZM-0926-SM-4DXSW
40-49 years, male
9.3
GTEX-UTHO-2726-SM-4JBH9
60-69 years, male
9.3
GTEX-17HGU-2226-SM-7IGLI
50-59 years, male
9.3
GTEX-1IDJD-2026-SM-A96TC
30-39 years, male
9.3
GTEX-QEG5-2426-SM-EYYVC
20-29 years, male
9.3
GTEX-1LG7Y-2426-SM-EVR57
50-59 years, male
9.2
GTEX-1OKEX-1826-SM-DTX8J
60-69 years, male
9.2
GTEX-11P82-1126-SM-5BC5K
20-29 years, male
9.1
GTEX-146FQ-1726-SM-5QGPX
30-39 years, male
9.1
GTEX-QMRM-1426-SM-CM2U2
50-59 years, male
9.1
GTEX-1CB4E-1726-SM-7IGMS
60-69 years, male
9.0
GTEX-WFON-1626-SM-4LVMV
40-49 years, male
9.0
GTEX-1KWVE-1226-SM-EV7AG
30-39 years, male
9.0
GTEX-1MGNQ-2226-SM-EV7A2
40-49 years, male
9.0
GTEX-Q2AI-1126-SM-48U19
40-49 years, male
8.9
GTEX-1S831-1226-SM-EVR5D
20-29 years, male
8.9
GTEX-XQ3S-2526-SM-4BOOG
20-29 years, male
8.8
GTEX-132QS-1126-SM-5P9GC
60-69 years, male
8.7
GTEX-14BIL-2126-SM-73KW3
50-59 years, male
8.7
GTEX-11P7K-0626-SM-5985Z
30-39 years, male
8.6
GTEX-11WQK-2726-SM-5EQMU
50-59 years, male
8.6
GTEX-13N2G-1826-SM-5KM1I
60-69 years, male
8.6
GTEX-147JS-2826-SM-5YYA7
60-69 years, male
8.6
GTEX-WY7C-2026-SM-4ONCM
50-59 years, male
8.6
GTEX-ZTX8-1026-SM-5EGHD
20-29 years, male
8.6
GTEX-16XZZ-2426-SM-7KULN
60-69 years, male
8.5
GTEX-NPJ8-2426-SM-3MJHL
40-49 years, male
8.5
GTEX-XAJ8-0626-SM-47JY4
40-49 years, male
8.5
GTEX-ZLFU-1726-SM-4WWBU
40-49 years, male
8.5
GTEX-1C6WA-2326-SM-7P8OW
60-69 years, male
8.5
GTEX-11ZUS-2126-SM-5N9E6
30-39 years, male
8.4
GTEX-17HII-2526-SM-7LT8C
50-59 years, male
8.4
GTEX-1B8KZ-1626-SM-73KUU
50-59 years, male
8.4
GTEX-SNOS-1026-SM-4DM4Y
40-49 years, male
8.4
GTEX-1J8Q3-0926-SM-A96SU
60-69 years, male
8.4
GTEX-1JKYN-0926-SM-AHZ47
30-39 years, male
8.4
GTEX-17HHY-2526-SM-7IGME
50-59 years, male
8.3
GTEX-P4QS-2026-SM-3NMCG
60-69 years, male
8.3
GTEX-RM2N-1126-SM-48FCY
50-59 years, male
8.3
GTEX-1EH9U-1826-SM-7P8QL
60-69 years, male
8.3
GTEX-OXRL-2026-SM-E6CHN
50-59 years, male
8.2
GTEX-131XF-5013-SM-7EWFI
60-69 years, male
8.1
GTEX-14ABY-0726-SM-5Q5DF
50-59 years, male
8.1
GTEX-16GPK-2826-SM-6LPJ6
60-69 years, male
8.1
GTEX-XGQ4-1826-SM-4AT6F
50-59 years, male
8.1
GTEX-XMK1-1026-SM-4B65H
40-49 years, male
8.1
GTEX-1GZ4I-2226-SM-9WPPO
50-59 years, male
8.1
GTEX-1I19N-2226-SM-A96TS
70-79 years, male
8.1
GTEX-1KAFJ-1226-SM-D4P46
50-59 years, male
8.1
GTEX-111YS-1726-SM-5GIED
60-69 years, male
8.0
GTEX-178AV-0426-SM-6LPJG
40-49 years, male
8.0
GTEX-1B8KE-1026-SM-731EQ
50-59 years, male
8.0
GTEX-1HKZK-1626-SM-B2LWH
40-49 years, male
8.0
GTEX-13O21-1126-SM-5N9EV
50-59 years, male
7.9
GTEX-1HSMO-2326-SM-CGQF3
60-69 years, male
7.9
GTEX-1IDJF-2126-SM-AHZ2S
20-29 years, male
7.9
GTEX-1JMPZ-2326-SM-CE6RW
30-39 years, male
7.9
GTEX-1JN6P-2626-SM-D4P3C
60-69 years, male
7.9
GTEX-1399T-1226-SM-5P9J5
60-69 years, male
7.8
GTEX-14PJ2-2126-SM-5YY96
50-59 years, male
7.8
GTEX-OOBJ-2026-SM-3NB1R
60-69 years, male
7.8
GTEX-WK11-2626-SM-4OOSK
50-59 years, male
7.8
GTEX-ZT9W-1626-SM-4YCDQ
50-59 years, male
7.8
GTEX-1JKYR-1726-SM-C1YR6
20-29 years, male
7.8
GTEX-1KANA-1026-SM-D3L9L
20-29 years, male
7.8
GTEX-1MJK2-0726-SM-DTXE7
50-59 years, male
7.8
GTEX-11EQ9-1726-SM-5HL6U
30-39 years, male
7.7
GTEX-13FTW-1226-SM-5LZZ1
40-49 years, male
7.7
GTEX-13O61-1726-SM-5KM46
60-69 years, male
7.7
GTEX-13RTJ-2126-SM-5S2PZ
60-69 years, male
7.7
GTEX-1EU9M-1526-SM-79OLI
60-69 years, male
7.7
GTEX-T5JC-1226-SM-4DM7C
20-29 years, male
7.7
GTEX-1ICLZ-2626-SM-A9G2K
60-69 years, male
7.7
GTEX-SIU8-2226-SM-EAZ2T
50-59 years, male
7.7
GTEX-12ZZZ-2026-SM-5LZWJ
60-69 years, male
7.6
GTEX-U3ZH-0926-SM-4DXU4
30-39 years, male
7.6
GTEX-ZTTD-2726-SM-57WFA
50-59 years, male
7.6
GTEX-1H3VY-2726-SM-A96TL
50-59 years, male
7.6
GTEX-1GZ2Q-2426-SM-9KNV8
40-49 years, male
7.5
GTEX-1OJC3-1926-SM-DPRYV
20-29 years, male
7.5
GTEX-QEG4-2226-SM-CXZJD
30-39 years, male
7.5
GTEX-QLQ7-1326-SM-CNNOR
60-69 years, male
7.5
GTEX-V955-1826-SM-4JBIL
60-69 years, male
7.4
GTEX-1IDJH-2426-SM-CNNQW
70-79 years, male
7.4
GTEX-OIZF-2026-SM-3MJH1
60-69 years, male
7.4
GTEX-WFG7-1826-SM-3GIL1
20-29 years, male
7.3
GTEX-1K2DA-1226-SM-CGQGH
50-59 years, male
7.3
GTEX-1QP6S-0726-SM-E76QS
40-49 years, male
7.3
GTEX-14BMV-2326-SM-5RQJ4
50-59 years, male
7.2
GTEX-1CB4G-2526-SM-7MXTG
60-69 years, male
7.2
GTEX-1E1VI-2726-SM-7MXVH
50-59 years, male
7.2
GTEX-1JJE9-0426-SM-D4P2P
70-79 years, male
7.2
GTEX-12BJ1-1226-SM-5LUAE
60-69 years, male
7.1
GTEX-17HHE-1126-SM-793BS
50-59 years, male
7.1
GTEX-1C6VQ-2426-SM-7EWEU
50-59 years, male
7.1
GTEX-ZVP2-0826-SM-59HJ6
50-59 years, male
7.1
GTEX-1F6I4-1326-SM-7P8QQ
60-69 years, male
7.1
GTEX-1H3NZ-2826-SM-9WPQ7
60-69 years, male
7.1
GTEX-1JMLX-1226-SM-CY8HW
50-59 years, male
7.1
GTEX-1LSNL-2326-SM-DHXKN
50-59 years, male
7.1
GTEX-11TT1-2026-SM-5EQM8
20-29 years, male
7.0
GTEX-ZYFD-2526-SM-5E45L
50-59 years, male
7.0
GTEX-1S5ZA-1626-SM-E9TKG
20-29 years, male
7.0
GTEX-1EKGG-2826-SM-7MGXA
60-69 years, male
6.8
GTEX-1GN2E-2026-SM-9JGGG
60-69 years, male
6.8
GTEX-1JK1U-0926-SM-AHZ43
60-69 years, male
6.8
GTEX-1L5NE-1026-SM-D3LAP
50-59 years, male
6.8
GTEX-14ICK-1826-SM-69LOP
40-49 years, male
6.7
GTEX-14PJ4-1226-SM-5YY9U
60-69 years, male
6.6
GTEX-15CHR-1226-SM-79OON
50-59 years, male
6.6
GTEX-1AX9K-1526-SM-73KVP
50-59 years, male
6.6
GTEX-1GF9V-0726-SM-9KNU4
60-69 years, male
6.6
GTEX-1HSMQ-1826-SM-A9SMJ
50-59 years, male
6.6
GTEX-13111-1326-SM-5GCNO
50-59 years, male
6.5
GTEX-14C5O-2826-SM-5RQI6
60-69 years, male
6.5
GTEX-14E6C-2826-SM-664NC
40-49 years, male
6.5
GTEX-14DAR-1026-SM-73KV3
50-59 years, male
6.4
GTEX-15RIF-1526-SM-6M46K
20-29 years, male
6.4
GTEX-OOBK-2025-SM-3LK5S
40-49 years, male
6.4
GTEX-14PJ3-1726-SM-5ZZVH
50-59 years, male
6.2
GTEX-XPVG-2026-SM-4B65E
50-59 years, male
6.2
GTEX-1AX9I-2326-SM-7PBXV
60-69 years, male
6.2
GTEX-1GL5R-1226-SM-7MXUY
50-59 years, male
6.2
GTEX-1JJ6O-0726-SM-CY8HS
60-69 years, male
6.2
GTEX-12696-2226-SM-5EQ65
60-69 years, male
6.1
GTEX-13OW8-0426-SM-5J2NR
60-69 years, male
6.1
GTEX-1BAJH-0626-SM-7P8OY
70-79 years, male
6.1
GTEX-S3XE-1326-SM-4AD4H
50-59 years, male
6.0
GTEX-1R7EU-0126-SM-EVR4F
60-69 years, male
6.0
GTEX-1AX9J-1426-SM-731BV
60-69 years, male
5.9
GTEX-1AX8Z-5019-SM-AHZ2C
60-69 years, male
5.9
GTEX-1GPI7-1626-SM-7P8TN
20-29 years, male
5.9
GTEX-1HFI6-2326-SM-9WYUN
40-49 years, male
5.9
GTEX-117YW-1426-SM-5EGGO
50-59 years, male
5.8
GTEX-13NYS-1026-SM-5KLZT
60-69 years, male
5.8
GTEX-14E6E-1126-SM-5S2R4
30-39 years, male
5.8
GTEX-1HSKV-1726-SM-C1YRO
60-69 years, male
5.8
GTEX-REY6-2126-SM-48FD9
60-69 years, male
5.7
GTEX-1PFEY-0626-SM-E76PH
30-39 years, male
5.7
GTEX-11O72-2726-SM-5HL6K
40-49 years, male
5.5
GTEX-1477Z-2226-SM-5QGPG
60-69 years, male
5.5
GTEX-ZG7Y-2226-SM-DO92I
50-59 years, male
5.5
GTEX-11DXZ-1826-SM-5H12Y
50-59 years, male
5.4
GTEX-ZEX8-2026-SM-4WKF3
50-59 years, male
5.4
GTEX-RVPV-2526-SM-EYYVL
60-69 years, male
5.3
GTEX-T6MN-2126-SM-EZ6LC
50-59 years, male
5.3
GTEX-XXEK-1226-SM-4BRUY
50-59 years, male
5.1
GTEX-1HBPN-2126-SM-9WPNQ
50-59 years, male
5.1
GTEX-SUCS-1126-SM-4DM61
70-79 years, male
5.0
GTEX-16MTA-1426-SM-6PALY
50-59 years, male
4.9
GTEX-SJXC-2026-SM-4DM6N
60-69 years, male
4.9
GTEX-1KXAM-1126-SM-E9TJU
60-69 years, male
4.9
GTEX-VJYA-1226-SM-3GIJ6
60-69 years, male
4.7
GTEX-1GN73-1626-SM-9WYTS
60-69 years, male
4.7
GTEX-145LU-2126-SM-5Q5E9
50-59 years, male
4.4
GTEX-U8XE-2526-SM-4E3IT
30-39 years, male
4.4
GTEX-1PWST-2626-SM-E8VN8
70-79 years, male
4.1
GTEX-1S3DN-1926-SM-E6CIR
50-59 years, male
4.1
GTEX-1GF9W-2026-SM-7MXVO
60-69 years, male
3.8
GTEX-1I6K7-2326-SM-ARZM2
40-49 years, male
3.8
Show allShow less
PROSTATE - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.